RM2AADK3X–Orange-finned anemonefish (Amphiprion chrysopterus) guarding their recently spawned red eggs on a rock next to the host anemone. Rock Islands, Palau, Micronesia. Tropical west Pacific Ocean.
RM2AADK3F–Yellowhead jawfish (Opistognathus aurifrons) spitting out sand as it digs its burrow. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea.
RM2AKEMBM–Striped mackerel (Rastrelliger kanagurta) shoal swimming with open mouths, filtering for zooplankton. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
RM2AKEM82–Raggy scorpionfish (Scorpaenopsis venosa) disputing territory. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean.
RM2AKEKRF–Florida manatee (Trichechus manatus latirostrus) with Blue gill sunfish (Lepomis macrochirus) cleaning it, in a freshwater spring, beneath trees. Three Sisters Spring, Crystal River, Florida, USA
RM2AADK39–Striped mackerel (Rastrelliger kanagurta) mouth wide open as it swims through the water, filtering zoopankton with its gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
RM2AADK3P–Mirrored portrait of a Weedy scorpionfish (Rhinopias frondosa). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea. Digitally manipulated (the face is reflected down mid-line).
RM2AKENCA–A threadfin hawkfish (Cirrhitichthys aprinus) on a barrel sponge. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
RM2AKENC6–Galapagos barnacle blenny (Acanthemblemaria castroi) looking out from its home in an old barnacle shell. Cape Marshall, Isabela Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean. Endemic.
RM2AADK38–Striped mackerel (Rastrelliger kanagurta) with mouths wide open as they swim through the water, filtering zoopankton with their gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
RM2AKEM74–Reticulate stargazer (Uranoscopus sp.) portrait. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean.
RM2AKEKPW–Schools of Pacific creolefish (Cephalopholis colonus) and Panamic sergeant major (Abudefduf troschelii) swimming through sun beams inside an underwater cavern. Los Islotes, La Paz, Baja California Sur, Mexico. Sea of Cortez, Gulf of California, East Pacific Ocean.
RM2AKEM7B–Giant mottled eel (Anguilla marmorata) in river with villagers washing clothes in background. Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean.
RM2AKEM7T–Oriental sweetlips (Plectorhinchus vittatus) pair. South Ari Atoll, Madlives. Indian Ocean.
RM2AADK43–Whitebelly damselfish (Amblyglyphidodon leucogaster) male and female laying egg on old gorgonian branch. The Alternatives, Sinai, Egypt. Red Sea.
RM2AADK41–Tiger cardinalfish (Cheilodipterus macrodon) male mouthbrooding eggs in his mouth. Gubal Barge, Gubal Island, Egypt. Strait of Gubal, Gulf of Suez, Red Sea
RM2AADK44–Bullethead parrotfish (Chlorurus sordidus) male, close up of eye, whilst sleeping at night. The Alternatives, Sinai, Egypt. Red Sea
RM2AKEKR2–Mandarinfish (Synchiropus splendidus) pair spawning. The larger male (behind) is lifting the female into the spawning rise on his pectoral fin, the smaller female (foreground) is releasing eggs, clearly visible below her body. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
RM2AADK3J–Latticetail moray (Gymnothorax buroensis) emerging from soft coral on a reef. Sangeang Island, Sumbawa, Indonesia. Flores Sea.
RM2AKEM7W–Bengal snapper (Lutjanus bengalensis), Bannerfish (Heniochus diphreutes) and Fusiliers (Caesio sp.) diving towards coral reef to avoid predators. North Ari Atoll, Maldives. Indian Ocean.
RM2AADK40–Bluestreak cleaner wrasse (Labroides dimidiatus) sticking its head into the mouth of a Red Sea fusilier (Caesio lunaris) during a cleaning session. Small Crack, Sha'ab Mahmud, Sinai, Egypt. Red Sea.
RMPBADNR–Fangtooth moray eel (Enchelycore anatina) with mouth open, Grand Canaria, Canary Islands, Spain. East Atlantic Ocean.
RMPBAGF5–Portrait of a male Squarespot anthias (Pseudanthias pleurotaenia). Sangeang Island; Sumbawa, Indonesia. Flores Sea.
RMPBAG0P–Ribbon sweetlips (Plectorhinchus polytaenia) USAT Liberty Wreck, Tulamben, Bali, Indonesia. Java Sea
RMPB8FM5–Warty / Clown frogfish (Antennarius maculatus) male portrait, Anilao, Batangas, Luzon, Philippines, March
RMPBAGF9–Spotlight goby (Gobiosoma louisae) looking out from inside a Yellow tube sponge (Aplysina fistularis), growing on a coral reef. Bloody Bay Wall, Little Cayman, Cayman Islands, British West Indies. Caribbean Sea.
RMPBADE4–Western clownfish (Amphiprion ocellaris) in front of an anemone, Tulamben Bay, Bali, Indonesia. Java Sea.
RMPBADE3–False clown anemonefish (Amphiprion ocellaris) large female in her host anemone, below rainforest. The Passage, between Gam and Waigeo Islands, West Papua, Indonesia. Composite image.
RMW7RF1T–Pair of golden rudd (Scardinius erythrophthalmus). Capernwray Lake, Lake District, Lancashire, England, UK.
RMW7REWD–Thermal pool filled with water that remains close to 30 degrees celsius all year, and is able to support a population of fancy goldfish (Carassius auratus) despite being close to the Arctic Circle. Husavik, Iceland. May 2011.
RMW7WNXN–School of Giant trevallies (Caranx ignobilis) in open water off the wall at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea.
RMW7WP2B–Shoal of Bar jacks (Caranx ruber) hunting school of Silversides fish (Atherinidae) in coral cavern. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea.
RM2ATWH6B–Whale shark (Rhincodon typus) close to the camera with a golden trevally fish (Gnathanodon speciosus) and outline of another whale shark behind , in the plankton-rich green waters of La Paz bay, Baja California Sur, Mexico. Sea of Cortez, Gulf of California, East Pacific Ocean.
RMW7YMD4–Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea, close to Little Brother Island. A rainbow runner (Elagatis bipinnulata) is hiding amonst the pilotfish. The Brothers Islands, Egypt. Red Sea. Endangered species.
RMGAATC1–Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea, close to Little Brother Island. The Brothers Islands, Egypt. Red Sea. Endangered species.
RMW7YMD1–Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea, close to Little Brother Island. The Brothers Islands, Egypt. Red Sea. Endangered species.
RM2AADGXA–Caribbean reef sharks (Carcharhinus perezi) swimming through the middle of a school of yellowtail snappers (Ocyurus chrysurus). Grand Bahama, Bahamas. Tropical West Atlantic Ocean.
RMM49FBR–Flame anthias (Pseudanthias ignitus) displaying to the more drab female (above) during courtship in the evening. Similan Islands, Thailand. Andaman Sea, Indian Ocean.
RMM49FG4–Leafy seadragon (Phycodurus eques). Wool Bay Jetty, Edithburgh, Yorke Peninsula, South Australia.
RMM49HED–Dwarf lionfish (Dendrochirus brachypterus) opens its ‘wings’ (in fact enlarged pectoral fins) Seraya, Tulamben area, Bali, Indonesia, Java Sea.
RMM49FG1–Nassau grouper (Epinephelus striatus) yawning on a coral reef drop off. San Salvador island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean.
RMM49FFJ–Longfin batfish (Platax teira) with Fusiliers (Caesio spp.) in the current in front of Baby Rock pinnacle. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea.
RMM49FBK–Wolf fish (Anarhichas lupus) portrait, Akuyeri, Iceland. North Atlantic Ocean
RMM49FFN–Bohar snappers (Lutjanus bohar) breaking up into smaller groups to spawn. The fish swim short arches above the main group as they release clouds of gametes. Shark City, Ulong, Rock Islands, Palau. Tropical Pacific Ocean.
RMM49HEJ–Silversides (Atherinomorus lacunosus) school just below surface in evening light, Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan.
RMM49HET–Crocodilefish (Cymbacephalus beauforti) close up detail of the retractable eye-lashes, Mabul Island, Sabah, Borneo, Malaysia Celebes Sea
RMM49HEE–Tompot blenny (Parablennius gattorugine) portrait, Swanage Pier, Swanage, Dorset, UK, English Channel, August
RMM49HF2–Bohar snapper (Lutjanus bohar) school at Ras Mohammed, Shark Reef, Ras Mohammed, Sinai, Egypt. Red Sea.
RMM494FX–Tompot blenny (Parablennius gattorugine) male peers out of its den, in leg of Swanage Pier, Swanage, Dorset, UK, English Channel, August
RMM49FFY–Brandt's cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig. Ellen Rig, Los Angeles, California, United States of America. North East Pacific Ocean.
RMM49GY2–Brandt's cormorant (Phalacrocorax penicillatus) hunts for a meal in a school of Pacific chub mackerel (Scomber japonicus), beneath an oil rig. Eureka Rig, Los Angeles, California, United States of America. North East Pacific Ocean. Finalist in the Birds Category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2016.
RMM49HEC–Bignose unicornfish (Naso vlamingii) profile of male displaying (the blue stripe at the bottom of the frame is the tail of another individual) Baa Atoll, Maldives. Indian Ocean.
RMM49HEB–California sea lion (Zalophus californianus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath Eureka oil rig, Los Angeles, California, USA, Pacific Ocean
RFHY1WD4–Mexican hogfish (Bodianus diplotaenia). Socorro Island, Revillagigedos, Mexico. East Pacific Ocean.
RFHY1X0N–High magnification photo of Secretary blenny (Acanthemblemaria maria) in boulder brain coral (Colpophyllia natans). East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea.
RFHY1X0C–Brandt's cormorant (Phalacrocorax penicillatus) hunting in school of Pacific chub mackerel (Scomber japonicus), beneath oil rig. Eureka Rig, Los Angeles, California, United States of America. North East Pacific Ocean.
RFHY1WYF–Shallow depth of field portrait of male Scalefin anthia (Pseudanthias squamipinnis) swimming in open water above coral reef photographed against sunset. The Alternatives, Sinai, Egypt. Gulf of Aqaba, Red Sea.
RFHY1WYD–Bluestreak cleaner wrasse (Labroides dimidiatus) tending to Ribboned sweetlips (Plectorhinchus polytaenia). Yillet Kecil, Yillet Islands, Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean. Ceram Sea.
RFHY1X0D–Pair of lionfish (Pterois volitans) prowling soft coral (Dendronephthya hemprichi) growing on wreck of the Cedar Pride, hunting glassfish. Aqaba, Jordan. Gulf of Aqaba, Red Sea.
RFHY1X09–Common ghost goby (Pleurosicya mossambica) on orange sponge. Aqaba, Jordan. Gulf of Aqaba, Red Sea.
RFHY1WYE–Young lionfish (Pterois volitans) swimming near surface hunting silversides, at dusk. Beacon Rock, Sha'ab Mahmood, Sinai, Egypt. Red Sea.
RFHY1WB3–Nassau grouper (Epinephelus striatus) with Cleaner gobies (Gobiosoma genie) and juvenile Spanish hogfish (Bodianus rufus) on coral reef. Bloody Bay Wall, Little Cayman, Cayman Islands. Caribbean Sea.
RFHY1WY9–Titan triggerfish (Balistoides viridescens) swimming over hard coral (Acropora sp.) gardens. Kri Island, Raja Ampat, West Papua, Indonesia. Dampier Strait, tropical west Pacific Ocean.
RFHY1TCT–Roughhead blenny (Acanthemblemaria aspera) female, high magnification of golden variety living in tube on coral reef. East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea.
RFHY1X10–A colourful reef scene with seafans (Melithaea sp.) soft corals (Dendronephthya sp) and mainly female scalefin anthias (Pseudanthias squamipinnis). Ra Province, Viti Levu, Fiji, Polynesia. Bligh Waters, Vatu-i-Ra Passage, Tropical South Pacific Ocean
RFHY1X11–School of large Atlantic bluefin tuna (Thunnus thynnus) captive in a growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea.
RFHY1T5C–Denise's Pygmy Seahorse (Hippocampus denise) on a seafan. Misool, Raja Ampat, West Papua, Indonesia.
RFHY1T5B–Orange Common / Spotted Seahorse (Hippocampus kuda) on seabed. Tulamben, Bali, Indonesia, Java Sea.
RFHY1TCH–Chain moray eel (Echidna catenata) portrait with mouth open. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea.
RM2BNEM5G–Andaman anemone fishes (Amphiprion ephippium) hovering above a sea anemone on a coral reef in Thailand. Richelieu Rock, Andaman Sea, Indian Ocean.
RM2BNEM63–School of Crescent-tail bigeyes (Priacanthis hamrur) swims in front of a school of Bigeye jacks (Caranx sexfasciatus). Ulong Island, Rock Islands, Pal
RM2BNH73Y–Green jacks (Caranx caballus) long exposure, La Paz, Mexico. Gulf of California, Sea of Cortez, East Pacific Ocean.
RM2BNH726–Diver photographing a school of Big eye trevally (Caranx sexfasciatus). South Atoll, Tubbataha Atolls, Tubbataha Reefs Natural Park, Palawan, Philippi
RM2BNM5T8–Bar jacks (Caranx ruber) pair hunting Silversides (Atherinidae) inside the wreck of the Kittiwake. Seven Mile Beach, Grand Cayman, Cayman Islands, Car
RM2BND533–Huge school of Bigeye trevally (Caranx sexfasciatus) swimming over a coral reef. South Atoll, Tubbataha Atolls, Tubbataha Reefs Natural Park, Palawan,
RM2BND531–Huge school of Bigeye trevally (Caranx sexfasciatus) swimming over a coral reef. South Atoll, Tubbataha Atolls, Tubbataha Reefs Natural Park, Palawan,
RM2BNM5JJ–Yellow striped scad (Selaroides leptolepis) school swim around a leg of Arborek jetty, Arborek Island, Raja Ampat, West Papua, Indonesia
RM2BNE02D–Oceanic whitetip shark (Carcharhinus longimanus) accompanied by a group of Pilotfish (Naucrates ductor). Rocky Island, Egypt. Red Sea
RM2BNEM5Y–Oceanic whitetip shark (Carcharhinus longimanus) cruising through water,. This individual is accompanied by a group of Pilotfish (Naucrates ductor), m
RM2BNM61E–Southern stingrays (Hypanus americanus) large female and two smaller males forage over seagrass in shallow water, accompanied by Bar jacks (Caranx rub
RM2BNEM5C–Great hammerhead shark (Sphyrna mokarran) swimming over a sandy seabed, accompanied by Rainbow runners (Elagatis bipinnulata) South Bimini, Bahamas. T
RM2BNH6X3–Slender grouper (Anyperodon leucogrammicus) exploiting the confusion caused by and Orange spotted trevally (Carangoides bajad) hunting silversides (At
RMW7RF1R–Loose shoal of young common roach (Rutilus rutilus) swimming amongst reeds, sedges and fallen trees in an English lake. Gildenburgh Water, Whittlesey, Cambridgeshire. Great Britain.
RMM49EP7–Oceanic whitetip shark (Carcharhinus longimanus) accompanied by a group of Pilotfish (Naucrates ductor). Rocky Island, Egypt. Red Sea
RMM49FBN–Great hammerhead shark (Sphyrna mokarran) female at night with Bar jacks (Caranx ruber), South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean.
RMM49FFX–School of Blacksmith fish (Chromis punctipinnis) swim through a giant kelp (Macrocystis pyrifera) forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacific Ocean.
RMM49HFC–Blacksmiths (Chromis punctipinnis) school swam through a giant kelp (Macrocystis pyrifera) forest, Santa Barbara Island, Channel Islands. Los Angeles, California, USA, North East Pacific Ocean.
RMM49HE5–Soft corals (Dendronephthya hemprichi) growing in very shallow water in the shade provided by a jetty, while a school of Silversides (Atherinomorus lacunosus) circle. Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan.
RMM49FC2–Orange anemonefish (Amphiprion sandaracinos) in its host Mertens' carpet sea anemone (Stichodactyla mertensii). Mabul Island, Sabah, Borneo, Malaysia. Celebes Sea / Sulawesi Sea.
RMG9XMAP–Long exposure of Black jack (Caranx lugubris) swimming through a cavern in a coral reef. East End, Grand Cayman, Cayman Islands.
RMG9XP27–Group of predators hunting together, in an example of 'Nuclear foraging'. The Moray eel (Gymnothorax javanicus) flushes out prey
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